About the Authors

Xin Jin

Contributed equally to this work with: Xin Jin, Mingze He, Betsy Ferguson, Yuhuan Meng, Limei Ouyang, Jingjing Ren

Affiliations School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China, BGI-Shenzhen, Shenzhen, China

Mingze He

Contributed equally to this work with: Xin Jin, Mingze He, Betsy Ferguson, Yuhuan Meng, Limei Ouyang, Jingjing Ren

Affiliations School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China, BGI-Shenzhen, Shenzhen, China

Betsy Ferguson

Contributed equally to this work with: Xin Jin, Mingze He, Betsy Ferguson, Yuhuan Meng, Limei Ouyang, Jingjing Ren

Affiliation Primate Genetics Program, Oregon National Primate Research Center, Oregon Health and Sciences University, Beaverton, Oregon, United States of America

Yuhuan Meng

Contributed equally to this work with: Xin Jin, Mingze He, Betsy Ferguson, Yuhuan Meng, Limei Ouyang, Jingjing Ren

Affiliation School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China

Limei Ouyang

Contributed equally to this work with: Xin Jin, Mingze He, Betsy Ferguson, Yuhuan Meng, Limei Ouyang, Jingjing Ren

Affiliation BGI-Shenzhen, Shenzhen, China

Jingjing Ren

Contributed equally to this work with: Xin Jin, Mingze He, Betsy Ferguson, Yuhuan Meng, Limei Ouyang, Jingjing Ren

Affiliations BGI-Shenzhen, Shenzhen, China, School of Biological Science and Medical Engineering, Southeast University, Nanjing, China

Thomas Mailund

Affiliation Bioinformatics Research Center, Aarhus University, Aarhus C, Denmark

Fei Sun

Affiliation School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China

Liangdan Sun

Affiliations Institute of Dermatology and Department of Dermatology, No.1 Hospital, Anhui Medical University, Hefei, Anhui, China, State Key Laboratory Incubation Base of Dermatology, Ministry of National Science and Technology, Hefei, Anhui, China

Juan Shen

Affiliation BGI-Shenzhen, Shenzhen, China

Min Zhuo

Affiliation School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China

Li Song

Affiliation BGI-Shenzhen, Shenzhen, China

Jufang Wang

Affiliation School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China

Fei Ling

Affiliation School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China

Yuqi Zhu

Affiliation BGI-Shenzhen, Shenzhen, China

Christina Hvilsom

Affiliations Science and Conservation, Copenhagen Zoo, Frederiksberg, Denmark, Bioinformatics, Department of Biology, University of Copenhagen, Copenhagen, Denmark

Hans Siegismund

Affiliation Bioinformatics, Department of Biology, University of Copenhagen, Copenhagen, Denmark

Xiaoming Liu

Affiliation South-China Primate Research and Development Center, Guangdong Entomological Institute, Guangzhou, China

Zhuolin Gong

Affiliation BGI-Shenzhen, Shenzhen, China

Fang Ji

Affiliation South-China Primate Research and Development Center, Guangdong Entomological Institute, Guangzhou, China

Xinzhong Wang

Affiliation South-China Primate Research and Development Center, Guangdong Entomological Institute, Guangzhou, China

Boqing Liu

Affiliation South-China Primate Research and Development Center, Guangdong Entomological Institute, Guangzhou, China

Yu Zhang

Affiliation Guangdong Laboratory Animals Monitoring Institute, Guangzhou, China

Jianguo Hou

Affiliation Guangdong Laboratory Animals Monitoring Institute, Guangzhou, China

Jing Wang

Affiliation Guangdong Laboratory Animals Monitoring Institute, Guangzhou, China

Hua Zhao

Affiliation Chinese PLA General Hospital, Beijing, China

Yanyi Wang

Affiliation Chinese PLA General Hospital, Beijing, China

Xiaodong Fang

Affiliation BGI-Shenzhen, Shenzhen, China

Guojie Zhang

Affiliation BGI-Shenzhen, Shenzhen, China

Jian Wang

Affiliation BGI-Shenzhen, Shenzhen, China

Xuejun Zhang

Affiliations Institute of Dermatology and Department of Dermatology, No.1 Hospital, Anhui Medical University, Hefei, Anhui, China, State Key Laboratory Incubation Base of Dermatology, Ministry of National Science and Technology, Hefei, Anhui, China

Mikkel H. Schierup

Affiliation Guangdong Laboratory Animals Monitoring Institute, Guangzhou, China

Hongli Du

hldu@scut.edu.cn (HD); wangj@genomics.cn (JW); xnwang@scut.edu.cn (XW)

Affiliation School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China

Jun Wang

hldu@scut.edu.cn (HD); wangj@genomics.cn (JW); xnwang@scut.edu.cn (XW)

Affiliations BGI-Shenzhen, Shenzhen, China, Bioinformatics, Department of Biology, University of Copenhagen, Copenhagen, Denmark

Xiaoning Wang

hldu@scut.edu.cn (HD); wangj@genomics.cn (JW); xnwang@scut.edu.cn (XW)

Affiliations School of Bioscience and Bioengineering, South China University of Technology, Guangzhou, China, Chinese PLA General Hospital, Beijing, China

Competing Interests

The authors have declared that no competing interests exist. Some of the authors are from Copenhagen Zoo, which is a non-profit organization and is listed as a cultural institution. In addition, Copenhagen Zoo has no commercial goals linked to the study, and this does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: XJ HD Xiaoning Wang Jian Wang Jun Wang HZ YW. Performed the experiments: BF FS L. Sun MZ Jufang Wang FL XL FJ Xinzhong Wang BL Y. Zhang JH Jing Wang XZ MHS. Analyzed the data: XJ MH BF YM LO JR JS L. Song Y. Zhu XF GZ. Wrote the paper: XJ BF TM MHS HD. Prepared samples: BF FS L. Sun MZ Jufang Wang FL XL FJ Xinzhong Wang BL Y. Zhang JH Jing Wang XZ MH. Assisted in writing manuscript: XJ MH BF YM LO JR TM FS L. Sun JS MZ L. Song Jufang Wang FL Y. Zhu CH HS XL ZG FJ Xinzhong Wang BL Yu Zhang JH Jing Wang HZ YW XF GZ Jian Wang XZ MHS HD Jun Wang Xiaoning Wang.