About the Authors

Dawne N. Shelton

Contributed equally to this work with: Dawne N. Shelton, Hubert Fornalik

Affiliation Department of Obstetrics and Gynecology, Holden Comprehensive Cancer Center, The University of Iowa Hospitals and Clinics, Iowa City, Iowa, United States of America

Hubert Fornalik

Contributed equally to this work with: Dawne N. Shelton, Hubert Fornalik

Affiliation Department of Obstetrics and Gynecology, Holden Comprehensive Cancer Center, The University of Iowa Hospitals and Clinics, Iowa City, Iowa, United States of America

Traci Neff

Affiliation Department of Obstetrics and Gynecology, Holden Comprehensive Cancer Center, The University of Iowa Hospitals and Clinics, Iowa City, Iowa, United States of America

Soo Yeun Park

Affiliation Department of Obstetrics and Gynecology, Holden Comprehensive Cancer Center, The University of Iowa Hospitals and Clinics, Iowa City, Iowa, United States of America

David Bender

Affiliation Department of Obstetrics and Gynecology, Holden Comprehensive Cancer Center, The University of Iowa Hospitals and Clinics, Iowa City, Iowa, United States of America

Koen DeGeest

Affiliation Department of Obstetrics and Gynecology, Holden Comprehensive Cancer Center, The University of Iowa Hospitals and Clinics, Iowa City, Iowa, United States of America

Xiaoming Liu

Affiliation Department of Anatomy and Cell Biology, The University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America

Weiliang Xie

Affiliation Department of Anatomy and Cell Biology, The University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America

David K. Meyerholz

Affiliation Department of Pathology, The University of Iowa Hospitals and Clinics, Iowa City, Iowa, United States of America

John F. Engelhardt

Affiliation Department of Anatomy and Cell Biology, The University of Iowa Carver College of Medicine, Iowa City, Iowa, United States of America

Michael J. Goodheart

michael-goodheart@uiowa.edu

Affiliation Department of Obstetrics and Gynecology, Holden Comprehensive Cancer Center, The University of Iowa Hospitals and Clinics, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DNS HF TN SYP JFE MJG. Performed the experiments: DNS HF TN SYP. Analyzed the data: DNS HF TN SYP XL WX DKM JFE MJG. Contributed reagents/materials/analysis tools: DB KD XL WX JFE MJG. Wrote the paper: DNS HF TN SYP DB KD DKM JFE MJG.