About the Authors

Marjo K. Hytönen

Affiliations Department of Veterinary Biosciences, Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, The Folkhälsan Institute of Genetics, Helsinki, Finland, Biochemistry and Developmental Biology, Institute of Biomedicine, University of Helsinki, Helsinki, Finland

Meharji Arumilli

Affiliations Department of Veterinary Biosciences, Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, The Folkhälsan Institute of Genetics, Helsinki, Finland

Anu K. Lappalainen

Affiliation Equine and Small Animal Medicine, Faculty of Veterinary Medicine, University of Helsinki, Helsinki, Finland

Heli Kallio

Affiliation Department of Veterinary Biosciences, Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland

Marjatta Snellman

Affiliation Equine and Small Animal Medicine, Faculty of Veterinary Medicine, University of Helsinki, Helsinki, Finland

Kirsi Sainio

Affiliation Biochemistry and Developmental Biology, Institute of Biomedicine, University of Helsinki, Helsinki, Finland

Hannes Lohi

hannes.lohi@helsinki.fi

Affiliations Department of Veterinary Biosciences, Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, The Folkhälsan Institute of Genetics, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MKH KS HL. Performed the experiments: MKH HK MS. Analyzed the data: MKH AKL HK MA KS HL. Contributed reagents/materials/analysis tools: MKH KS HL. Wrote the paper: MKH HL.