About the Authors

Kai He

Contributed equally to this work with: Kai He, Jian-Hai Chen, Gina C. Gould

Affiliations State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China, Graduate University of Chinese Academy of Sciences, Beijing, China

Jian-Hai Chen

Contributed equally to this work with: Kai He, Jian-Hai Chen, Gina C. Gould

Affiliations State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China, Graduate University of Chinese Academy of Sciences, Beijing, China

Gina C. Gould

Contributed equally to this work with: Kai He, Jian-Hai Chen, Gina C. Gould

Affiliations University of Connecticut, Department of Ecology and Evolutionary Biology, Stamford, Connecticut, United States of America, Bruce Museum, Greenwich, Connecticut, United States of America

Nobuyuki Yamaguchi

Affiliation Department of Biological & Environmental Sciences, Qatar University, Doha, Qatar

Huai-Sen Ai

Affiliation Baoshan Administrative Bureau, Gaoligong National Nature Reserve, Baoshan, China

Ying-Xiang Wang

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Ya-Ping Zhang

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Xue-Long Jiang

jiangxl@mail.kiz.ac.cn

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XLJ. Performed the experiments: KH JHC. Analyzed the data: KH JHC GCG. Contributed reagents/materials/analysis tools: KH JHC GCG NY HSA YXW YPZ. Wrote the paper: KH JHC GCG. Collected and examined the morphological characters: GCG.