About the Authors

Justin J. Greenlee

justin.greenlee@ars.usda.gov

Affiliation National Animal Disease Center, United States Department of Agriculture, Agricultural Research Service, Ames, Iowa, United States of America

Jodi D. Smith

Affiliation National Animal Disease Center, United States Department of Agriculture, Agricultural Research Service, Ames, Iowa, United States of America

M. Heather West Greenlee

Affiliation Iowa State University, Ames, Iowa, United States of America

Eric M. Nicholson

Affiliation National Animal Disease Center, United States Department of Agriculture, Agricultural Research Service, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJG JDS MHWG EMN. Performed the experiments: JJG JDS MHWG EMN. Analyzed the data: JJG JDS MHWG EMN. Contributed reagents/materials/analysis tools: JJG EMN. Wrote the paper: JJG JDS MHWG EMN.