About the Authors

Sei Higuchi

Affiliation Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan

Keiichi Irie

Affiliations Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan, Institute for Aging and Brain Sciences, Fukuoka University, Fukuoka, Japan

Ryuji Yamaguchi

Affiliation Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan

Mai Katsuki

Affiliation Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan

Maiko Araki

Affiliation Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan

Makiko Ohji

Affiliation Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan

Kazuhide Hayakawa

Affiliation Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan

Shohei Mishima

Affiliation Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan

Yoshiharu Akitake

Affiliations Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan, Institute for Aging and Brain Sciences, Fukuoka University, Fukuoka, Japan

Kiyoshi Matsuyama

Affiliation Department of Chemical Engineering, Faculty of Engineering, Fukuoka University, Fukuoka, Japan

Kenji Mishima

Affiliation Department of Chemical Engineering, Faculty of Engineering, Fukuoka University, Fukuoka, Japan

Kenichi Mishima

kenichi@fukuoka-u.ac.jp

Affiliations Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan, Institute for Aging and Brain Sciences, Fukuoka University, Fukuoka, Japan

Katsunori Iwasaki

Affiliations Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan, Institute for Aging and Brain Sciences, Fukuoka University, Fukuoka, Japan

Michihiro Fujiwara

Affiliation Department of Neuropharmacology, Faculty of Pharmaceutical Sciences, Fukuoka University, Fukuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SH YA K. Irie KH K. Mishima K. Iwasaki MF. Performed the experiments: SH MK RY MA MO SM. Analyzed the data: SH K. Irie K. Mishima MF. Contributed reagents/materials/analysis tools: K. Matsuyama K. Mishima MF. Wrote the paper: SH.