About the Authors

Wei Zhang

Contributed equally to this work with: Wei Zhang, John Ferguson

Affiliation Department of Medicine, Section of Digestive Diseases, Yale University School of Medicine, New Haven, Connecticut, United States of America

John Ferguson

Contributed equally to this work with: Wei Zhang, John Ferguson

Affiliation Department of Medicine, Section of Digestive Diseases, Yale University School of Medicine, New Haven, Connecticut, United States of America

Sok Meng Ng

Affiliation Department of Medicine, Section of Digestive Diseases, Yale University School of Medicine, New Haven, Connecticut, United States of America

Ken Hui

Affiliation Department of Medicine, Section of Digestive Diseases, Yale University School of Medicine, New Haven, Connecticut, United States of America

Gerald Goh

Affiliation Department of Genetics, Yale University School of Medicine, New Haven, Connecticut, United States of America

Aiping Lin

Affiliation W.M. Keck Biotechnology Laboratory, Yale University School of Medicine, New Haven, Connecticut, United States of America

Enric Esplugues

Affiliation Department of Immunobiology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Richard A. Flavell

Affiliations Department of Immunobiology, Yale University School of Medicine, New Haven, Connecticut, United States of America, Howard Hughes Medical Institute, Yale University School of Medicine, New Haven, Connecticut, United States of America

Clara Abraham

Affiliation Department of Medicine, Section of Digestive Diseases, Yale University School of Medicine, New Haven, Connecticut, United States of America

Hongyu Zhao

Affiliations Department of Genetics, Yale University School of Medicine, New Haven, Connecticut, United States of America, Department of Epidemiology and Public Health, Yale School of Medicine, New Haven, Connecticut, United States of America, Department of Statistics, Yale University School of Medicine, New Haven, Connecticut, United States of America

Judy H. Cho

judy.cho@yale.edu

Affiliations Department of Medicine, Section of Digestive Diseases, Yale University School of Medicine, New Haven, Connecticut, United States of America, Department of Genetics, Yale University School of Medicine, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WZ JF EE RAF CA HZ JHC. Performed the experiments: WZ SN. Analyzed the data: WZ JF AL KH GG JHC. Wrote the paper: WZ JF JHC.