About the Authors

Patricia J. Birkett

birkett.patricia@gmail.com

Affiliation Amarula Elephant Research Programme, School of Life Sciences, University of Kwazulu-Natal, Durban, South Africa

Abi T. Vanak

Current address: Ashoka Trust for Research in Ecology and the Environment. Bangalore, India

Affiliation Amarula Elephant Research Programme, School of Life Sciences, University of Kwazulu-Natal, Durban, South Africa

Vito M. R. Muggeo

Affiliation Dipartimento di Scienze Statistiche e Matematiche ‘Vianelli’, Università di Palermo, Palermo, Italy

Salamon M. Ferreira

Affiliations Amarula Elephant Research Programme, School of Life Sciences, University of Kwazulu-Natal, Durban, South Africa, Scientific Services, Kruger National Park, Skukuza, South Africa

Rob Slotow

Affiliation Amarula Elephant Research Programme, School of Life Sciences, University of Kwazulu-Natal, Durban, South Africa

Competing Interests

Dr. Sam Ferreira is a research manager and employee of SANParks Scientific Services, Kruger National Park. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Analyzed the data: PJB ATV VMRM. Wrote the paper: PJB. Conceived and designed the research questions and analysis: PJB ATV VMRM SMF RS. Provided supervisory inputs and support: RS SMF. Provided inputs to manuscript: ATV VMRM SMF RS.