About the Authors

Samuel Zamora

Affiliation Department of Palaeontology, The Natural History Museum, London, United Kingdom

Imran A. Rahman

Affiliation School of Geography, Earth & Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, United Kingdom

Andrew B. Smith

a.smith@nhm.ac.uk

Affiliation Department of Palaeontology, The Natural History Museum, London, United Kingdom

Competing Interests

The authors received financial support for excavations from the bank Caja de Ahorros de la Inmaculada, Aragon, Spain. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Analyzed the data: SZ IAR. Wrote the paper: ABS SZ. Contributed equally to the description and interpretation of the fossils: SZ IAR ABS.