About the Authors

Jin Tian

Contributed equally to this work with: Jin Tian, Wenbao Qi

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangzhou, P. R. China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, P. R. China

Wenbao Qi

Contributed equally to this work with: Jin Tian, Wenbao Qi

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangzhou, P. R. China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, P. R. China

Xiaokang Li

Affiliation College of Animal Science and Technology, Henan University of Science and Technology, Luoyang, Henan, P. R. China

Jun He

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangzhou, P. R. China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, P. R. China

Peirong Jiao

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangzhou, P. R. China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, P. R. China

Changhui Zhang

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangzhou, P. R. China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, P. R. China

Guo-Qian Liu

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangzhou, P. R. China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, P. R. China

Ming Liao

mliao@scau.edu.cn

Affiliations College of Veterinary Medicine, South China Agricultural University, Guangzhou, P. R. China, MOA Key Laboratory for Animal Vaccine Development, Guangzhou, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JT WQ ML. Performed the experiments: JT CZ. Analyzed the data: JT XL JH. Contributed reagents/materials/analysis tools: GQL PJ. Wrote the paper: JT.