About the Authors

Evropi Theodoratou

E.Theodoratou@ed.ac.uk

Affiliations Centre for Population Health Sciences, University of Edinburgh, Edinburgh, United Kingdom, MRC Human Genetics Unit, Colon Cancer Genetics Group and Academic Coloproctology, Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital Edinburgh, United Kingdom

Tom Palmer

Affiliation MRC Centre for Causal Analyses in Translational Epidemiology, School of Social and Community Medicine, University of Bristol, Bristol, United Kingdom

Lina Zgaga

Affiliations Centre for Population Health Sciences, University of Edinburgh, Edinburgh, United Kingdom, MRC Human Genetics Unit, Colon Cancer Genetics Group and Academic Coloproctology, Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital Edinburgh, United Kingdom, Andrija Stampar School of Public Health, Medical School, University of Zagreb, Zagreb, Croatia

Susan M. Farrington

Affiliation MRC Human Genetics Unit, Colon Cancer Genetics Group and Academic Coloproctology, Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital Edinburgh, United Kingdom

Paul McKeigue

Affiliation Centre for Population Health Sciences, University of Edinburgh, Edinburgh, United Kingdom

Farhat V. N. Din

Affiliation MRC Human Genetics Unit, Colon Cancer Genetics Group and Academic Coloproctology, Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital Edinburgh, United Kingdom

Albert Tenesa

Affiliations MRC Human Genetics Unit, Colon Cancer Genetics Group and Academic Coloproctology, Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital Edinburgh, United Kingdom, The Roslin Institute, The University of Edinburgh, Easter Bush, Roslin, Scotland

George Davey-Smith

Affiliation MRC Centre for Causal Analyses in Translational Epidemiology, School of Social and Community Medicine, University of Bristol, Bristol, United Kingdom

Malcolm G. Dunlop

Affiliation MRC Human Genetics Unit, Colon Cancer Genetics Group and Academic Coloproctology, Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital Edinburgh, United Kingdom

Harry Campbell

Affiliation Centre for Population Health Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ET HC GDS PM MD. Performed the experiments: ET TP. Analyzed the data: ET. Contributed reagents/materials/analysis tools: TP LZ SMF PM FVND AT MD. Wrote the paper: ET TP LZ.