About the Authors

Andrew K. MacLeod

Contributed equally to this work with: Andrew K. MacLeod, Gail Davies

andrew.macleod@ed.ac.uk (AKM); i.deary@ed.ac.uk (IJD)

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Medical Genetics Section, Centre for Molecular Medicine, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Gail Davies

Contributed equally to this work with: Andrew K. MacLeod, Gail Davies

Affiliation Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Antony Payton

Affiliation Centre for Integrated Genomic Medical Research, University of Manchester, Manchester, United Kingdom

Albert Tenesa

Affiliations The Roslin Institute, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Edinburgh, United Kingdom, Medical Research Council Human Genetics Unit, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Sarah E. Harris

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Medical Genetics Section, Centre for Molecular Medicine, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

David Liewald

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Xiayi Ke

Affiliation Medical Research Council Centre of Epidemiology for Child Health, University College London Institute of Child Health, London, United Kingdom

Michelle Luciano

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Lorna M. Lopez

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Alan J. Gow

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Janie Corley

Affiliation Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Paul Redmond

Affiliation Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Geraldine McNeill

Affiliation Institute of Applied Health Sciences, University of Aberdeen, Aberdeen, United Kingdom

Andrew Pickles

Affiliation School of Epidemiology and Health Science, Department of Medicine, University of Manchester, Manchester, United Kingdom

William Ollier

Affiliation Centre for Integrated Genomic Medical Research, University of Manchester, Manchester, United Kingdom

Michael Horan

Affiliation School of Community-Based Medicine, Neurodegeneration Research Group, University of Manchester, Manchester, United Kingdom

John M. Starr

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Geriatric Medicine Unit, University of Edinburgh, Edinburgh, United Kingdom

Neil Pendleton

Affiliation School of Community-Based Medicine, Neurodegeneration Research Group, University of Manchester, Manchester, United Kingdom

Pippa A. Thomson

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Medical Genetics Section, Centre for Molecular Medicine, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

David J. Porteous

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Medical Genetics Section, Centre for Molecular Medicine, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Ian J. Deary

andrew.macleod@ed.ac.uk (AKM); i.deary@ed.ac.uk (IJD)

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IJD AT NP. Analyzed the data: AKM GD ML A. Payton. Contributed reagents/materials/analysis tools: SEH A. Payton ML LML AJG JC PR LJW GMcN DJP MH JMS NP IJD XK WO A. Pickles. Wrote the paper: AKM IJD DJP PAT. Performed quality control analyses and prepared data: AKM GD SEH DL AT ML LML Contributed to discussions regarding analyses: IJD DJP PAT ML.