About the Authors

Yongjun Fang

Contributed equally to this work with: Yongjun Fang, Hao Wu, Tongwu Zhang

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, James D. Watson Institute of Genome Sciences, Zhejiang University, Hangzhou, China, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Hao Wu

Contributed equally to this work with: Yongjun Fang, Hao Wu, Tongwu Zhang

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, James D. Watson Institute of Genome Sciences, Zhejiang University, Hangzhou, China, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Tongwu Zhang

Contributed equally to this work with: Yongjun Fang, Hao Wu, Tongwu Zhang

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, James D. Watson Institute of Genome Sciences, Zhejiang University, Hangzhou, China, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Meng Yang

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Yuxin Yin

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Linlin Pan

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Xiaoguang Yu

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Xiaowei Zhang

junyu@big.ac.cn (JY); zhangxw@big.ac.cn (XZ); husn@big.ac.cn (SH); imssallem@kacst.edu.sa (ISAM)

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Songnian Hu

junyu@big.ac.cn (JY); zhangxw@big.ac.cn (XZ); husn@big.ac.cn (SH); imssallem@kacst.edu.sa (ISAM)

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, James D. Watson Institute of Genome Sciences, Zhejiang University, Hangzhou, China, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Ibrahim S. Al-Mssallem

junyu@big.ac.cn (JY); zhangxw@big.ac.cn (XZ); husn@big.ac.cn (SH); imssallem@kacst.edu.sa (ISAM)

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China, Department of Biotechnology, College of Agriculture and Food Sciences, King Faisal University, Hofuf, Kingdom of Saudi Arabia

Jun Yu

junyu@big.ac.cn (JY); zhangxw@big.ac.cn (XZ); husn@big.ac.cn (SH); imssallem@kacst.edu.sa (ISAM)

Affiliations Joint Center for Genomics Research (JCGR), King Abdulaziz City for Science and Technology (KACST) and Chinese Academy of Sciences (CAS), Riyadh, Kingdom of Saudi Arabia, James D. Watson Institute of Genome Sciences, Zhejiang University, Hangzhou, China, CAS Key Laboratory of Genome Sciences and Information, Beijing Institute of Genomics (BIG), Chinese Academy of Sciences (CAS), Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XZ SH ISAM JY. Performed the experiments: YF YY LP. Analyzed the data: YF HW TZ. Contributed reagents/materials/analysis tools: MY XY. Wrote the paper: YF HW TZ.