About the Authors

Simone Mäntele

Affiliation Faculty of Health and Medical Sciences, University of Surrey, Guildford, United Kingdom

Daniella T. Otway

Affiliation Faculty of Health and Medical Sciences, University of Surrey, Guildford, United Kingdom

Benita Middleton

Affiliation Faculty of Health and Medical Sciences, University of Surrey, Guildford, United Kingdom

Silvia Bretschneider

Affiliation Faculty of Health and Medical Sciences, University of Surrey, Guildford, United Kingdom

John Wright

Affiliation Faculty of Health and Medical Sciences, University of Surrey, Guildford, United Kingdom

M. Denise Robertson

Affiliation Faculty of Health and Medical Sciences, University of Surrey, Guildford, United Kingdom

Debra J. Skene

Affiliation Faculty of Health and Medical Sciences, University of Surrey, Guildford, United Kingdom

Jonathan D. Johnston

j.johnston@surrey.ac.uk

Affiliation Faculty of Health and Medical Sciences, University of Surrey, Guildford, United Kingdom

Competing Interests

Partial funding was received from a commercial source (Stockgrand Ltd., UK). DJS and BM are co-directors of Stockgrand Ltd., UK. However, their involvement in the study was purely through their academic positions. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JDJ DJS MDR. Performed the experiments: SM DTO BM SB JW. Analyzed the data: SM DTO BM SB JW MDR DJS JDJ. Wrote the paper: JDJ SM.