About the Authors

Sanzhen Liu

Affiliation Department of Agronomy, Iowa State University, Ames, Iowa, United States of America

Cheng-Ting Yeh

Affiliation Department of Agronomy, Iowa State University, Ames, Iowa, United States of America

Ho Man Tang

Affiliation Interdepartmental Genetics Graduate Program, Iowa State University, Ames, Iowa, United States of America

Dan Nettleton

Affiliation Department of Statistics, Iowa State University, Ames, Iowa, United States of America

Patrick S. Schnable

schnable@iastate.edu

Affiliations Department of Agronomy, Iowa State University, Ames, Iowa, United States of America, Center for Plant Genomics, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SL PSS DN. Performed the experiments: SL HT. Analyzed the data: SLL CTY DN. Contributed reagents/materials/analysis tools: PSS. Wrote the paper: SL DT PSS.