About the Authors

Debra A. Thompson

Contributed equally to this work with: Debra A. Thompson, Naheed W. Khan, Mohammad I. Othman

Affiliations Department of Ophthalmology and Visual Sciences, University of Michigan Medical School, Ann Arbor, Michigan, United States of America, Department of Biological Chemistry, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Naheed W. Khan

Contributed equally to this work with: Debra A. Thompson, Naheed W. Khan, Mohammad I. Othman

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Mohammad I. Othman

Contributed equally to this work with: Debra A. Thompson, Naheed W. Khan, Mohammad I. Othman

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Bo Chang

Affiliation Jackson Laboratory, Bar Harbor, Maine, United States of America

Lin Jia

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Garrett Grahek

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Zhijian Wu

Affiliation Neurobiology-Neurodegeneration & Repair laboratory, National Eye Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Suja Hiriyanna

Affiliation Neurobiology-Neurodegeneration & Repair laboratory, National Eye Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Jacob Nellissery

Affiliation Neurobiology-Neurodegeneration & Repair laboratory, National Eye Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Tiansen Li

Affiliation Neurobiology-Neurodegeneration & Repair laboratory, National Eye Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Hemant Khanna

Affiliation Department of Ophthalmology, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Peter Colosi

Affiliation Neurobiology-Neurodegeneration & Repair laboratory, National Eye Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Anand Swaroop

swaroopa@nei.nih.gov (AS); jrheck@umich.edu (JRH)

Affiliation Neurobiology-Neurodegeneration & Repair laboratory, National Eye Institute, National Institutes of Health, Bethesda, Maryland, United States of America

John R. Heckenlively

swaroopa@nei.nih.gov (AS); jrheck@umich.edu (JRH)

Affiliation Department of Ophthalmology and Visual Sciences, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DAT NWK MIO BC AS JRH. Performed the experiments: NWK MIO BC LJ GG ZW SH JN. Analyzed the data: DAT NWK BC TL HK PC AS JRH. Contributed reagents/materials/analysis tools: DAT BC TL AS JRH. Wrote the paper: DAT NWK MIO HK PC AS JRH.