About the Authors

Nicholas A. Bokulich

Affiliations Department of Viticulture and Enology, Robert Mondavi Institute of Wine and Food Science, University of California Davis, Davis, California, United States of America, Department of Food Science and Technology, Robert Mondavi Institute of Wine and Food Science, University of California Davis, Davis, California, United States of America

Charles W. Bamforth

Affiliation Department of Food Science and Technology, Robert Mondavi Institute of Wine and Food Science, University of California Davis, Davis, California, United States of America

David A. Mills

damills@ucdavis.edu

Affiliations Department of Viticulture and Enology, Robert Mondavi Institute of Wine and Food Science, University of California Davis, Davis, California, United States of America, Department of Food Science and Technology, Robert Mondavi Institute of Wine and Food Science, University of California Davis, Davis, California, United States of America

Competing Interests

NB has received multiple scholarships donated by commercial funding sources (Briess Malt, Cargill Malt, Wine spectator; see below). All scholarships were reviewed and awarded by third-party sources (American Society of Brewing Chemists, UC Davis Department of Viticulture and Enology) and none of these companies had any contact with the authors. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DAM NAB CWB. Performed the experiments: NAB. Analyzed the data: DAM NAB. Contributed reagents/materials/analysis tools: DAM CWB. Wrote the paper: NAB DAM CWB.