About the Authors

Deborah I. Ritter

Current address: The Human Genome Sequencing Center, Baylor College of Medicine, Houston, Texas, United States of America

Affiliation Department of Biology, Boston College, Chestnut Hill, Massachusetts, United States of America

Zhiqiang Dong

Affiliation Department of Bioengineering and Therapeutic Sciences, Programs in Biological Sciences and Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Su Guo

chuangj@bc.edu (JHC); Su.Guo@ucsf.edu (SG)

Affiliation Department of Bioengineering and Therapeutic Sciences, Programs in Biological Sciences and Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Jeffrey H. Chuang

chuangj@bc.edu (JHC); Su.Guo@ucsf.edu (SG)

Affiliation Department of Biology, Boston College, Chestnut Hill, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Edited final manuscript: DIR ZD SG JHC. Manuscript Figures: DIR JHC. Designed and built website: DIR. Conceived and designed the experiments: DIR SG JHC. Performed the experiments: DIR ZD. Analyzed the data: DIR ZD SG JHC. Contributed reagents/materials/analysis tools: DIR ZD SG JHC. Wrote the paper: DIR JHC SG.