About the Authors

Kai Yu Zhang

Affiliation Department of Bioengineering, Imperial College London, South Kensington, London, United Kingdom

Alexis Wiktorowicz-Conroy

Affiliation Structure and Motion Laboratory, The Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

John R. Hutchinson

Affiliation Structure and Motion Laboratory, The Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Michael Doube

Affiliations Department of Bioengineering, Imperial College London, South Kensington, London, United Kingdom, Max Planck Institute for Molecular Cell Biology and Genetics, Dresden, Germany

Michal Klosowski

Affiliation Department of Bioengineering, Imperial College London, South Kensington, London, United Kingdom

Sandra J. Shefelbine

Affiliation Department of Bioengineering, Imperial College London, South Kensington, London, United Kingdom

Anthony M. J. Bull

a.bull@imperial.ac.uk

Affiliation Department of Bioengineering, Imperial College London, South Kensington, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AMJB KYZ. Performed the experiments: KYZ. Analyzed the data: KYZ AMJB. Contributed reagents/materials/analysis tools: KYZ MD MK AWC JRH. Wrote the paper: KYZ AWC AMJB JRH SJS MD.