About the Authors

Kathryn E. Marshall

Affiliation Marine Biotechnology, Marine Sciences Laboratory, Pacific Northwest National Laboratory, Sequim, Washington, United States of America

Errol W. Robinson

Affiliation Environmental Molecular Sciences Laboratory, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Shawna M. Hengel

Affiliation Environmental Molecular Sciences Laboratory, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Ljiljana Paša-Tolić

Affiliation Environmental Molecular Sciences Laboratory, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Guritno Roesijadi

g.roesijadi@pnnl.gov

Affiliation Marine Biotechnology, Marine Sciences Laboratory, Pacific Northwest National Laboratory, Sequim, Washington, United States of America

Competing Interests

The authors have no support or funding to report.

Author Contributions

Conceived and designed the experiments: KEM GR. Performed the experiments: KEM EWR SMH. Analyzed the data: KEM EWR SMH LPT GR. Contributed reagents/materials/analysis tools: KEM EWR LPT GR. Wrote the paper: KEM GR. Reviewed and edited the manuscript: EWR LPT.