About the Authors

Wendy Atkinson

Contributed equally to this work with: Wendy Atkinson, Reza Forghani

Affiliation Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America

Reza Forghani

Contributed equally to this work with: Wendy Atkinson, Reza Forghani

Affiliations Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America, Division of Neuroradiology, Department of Radiology, Massachusetts General Hospital and Harvard Medical School, Boston, Massachusetts, United States of America, Sir Mortimer B. Davis Jewish General Hospital and McGill University, Montréal, Quebec, Canada

Gregory R. Wojtkiewicz

Affiliation Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America

Benjamin Pulli

Affiliation Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America

Yoshiko Iwamoto

Affiliation Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America

Takuya Ueno

Affiliation Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America

Peter Waterman

Affiliation Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America

Jessica Truelove

Affiliation Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America

Rahmi Oklu

Affiliation Division of Vascular Imaging and Intervention, Department of Radiology, Massachusetts General Hospital and Harvard Medical School, Boston, Massachusetts, United States of America

John W. Chen

jwchen@mgh.harvard.edu

Affiliations Center for Systems Biology, Massachusetts General Hospital and Harvard Medical School, Richard B. Simches Research Center, Boston, Massachusetts, United States of America, Division of Neuroradiology, Department of Radiology, Massachusetts General Hospital and Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WA RF BP TU RO JWC. Performed the experiments: WA RF GRW BP YI TU PW JT. Analyzed the data: WA RF GRW BP PW JT RO JWC. Contributed reagents/materials/analysis tools: PW JT TU. Wrote the paper: WA RF GRW BP YI TU PW JT RO JWC.