About the Authors

Minako Izutsu

Contributed equally to this work with: Minako Izutsu, Naoyuki Fuse

Affiliations Laboratory for Biodiversity, Global COE Program, Graduate School of Science, Kyoto University, Kyoto, Japan, Laboratory for Molecular Developmental Biology, Graduate School of Science, Kyoto University, Kyoto, Japan

Jun Zhou

Affiliation Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America

Yuzo Sugiyama

Affiliation Laboratory for Biodiversity, Global COE Program, Graduate School of Science, Kyoto University, Kyoto, Japan

Osamu Nishimura

Affiliation Laboratory for Biodiversity, Global COE Program, Graduate School of Science, Kyoto University, Kyoto, Japan

Tomoyuki Aizu

Affiliation Comparative Genomics Laboratory, National Institute of Genetics, Mishima, Japan

Atsushi Toyoda

Affiliation Comparative Genomics Laboratory, National Institute of Genetics, Mishima, Japan

Asao Fujiyama

Affiliation Comparative Genomics Laboratory, National Institute of Genetics, Mishima, Japan

Kiyokazu Agata

Affiliations Laboratory for Biodiversity, Global COE Program, Graduate School of Science, Kyoto University, Kyoto, Japan, Laboratory for Molecular Developmental Biology, Graduate School of Science, Kyoto University, Kyoto, Japan

Naoyuki Fuse

Contributed equally to this work with: Minako Izutsu, Naoyuki Fuse

nfuse@gcoe.biol.sci.kyoto-u.ac.jp

Affiliation Laboratory for Biodiversity, Global COE Program, Graduate School of Science, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NF KA AF. Performed the experiments: MI JZ YS TA NF. Analyzed the data: MI JZ ON TA AT NF. Contributed reagents/materials/analysis tools: JZ ON AT AT KA NF. Wrote the paper: NF MI JZ.