About the Authors

Erin D. Scully

Affiliation Intercollege Program in Genetics, The Huck Institutes of the Life Sciences, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Kelli Hoover

Affiliation Department of Entomology, Center for Chemical Ecology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

John Carlson

Affiliations School of Forest Resources, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Department of Bioenergy Science and Technology (World Class University), Chonnam National University, Buk-Gu, Gwangju, Korea

Ming Tien

Affiliation Department of Biochemistry and Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Scott M. Geib

scott.geib@ars.usda.gov

Affiliation Tropical Crop and Commodity Protection Research Unit, USDA-ARS Pacific Basin Agricultural Research Center, Hilo, Hawaii, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SMG KH MT JC. Performed the experiments: SMG EDS. Analyzed the data: SMG EDS. Contributed reagents/materials/analysis tools: SMG KH JC MT. Wrote the paper: SMG KH EDS MT JC.