About the Authors

Matthew L. Farnsworth

matt.farnsworth@aphis.usda.gov

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Veterinary Service, Centers for Epidemiology and Animal Health, Fort Collins, Colorado, United States of America

Ryan S. Miller

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Veterinary Service, Centers for Epidemiology and Animal Health, Fort Collins, Colorado, United States of America

Kerri Pedersen

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Disease Program, Fort Collins, Colorado, United States of America

Mark W. Lutman

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Disease Program, Fort Collins, Colorado, United States of America

Seth R. Swafford

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Disease Program, Fort Collins, Colorado, United States of America

Philip D. Riggs

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Veterinary Service, Centers for Epidemiology and Animal Health, Fort Collins, Colorado, United States of America

Colleen T. Webb

Affiliation Department of Biology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MLF RSM CTW. Performed the experiments: MLF RSM. Analyzed the data: MLF RSM. Contributed reagents/materials/analysis tools: MLF RSM KP MWL SRS PDR. Wrote the paper: MLF RSM KP MWL SRS.