About the Authors

Wael Osman

Affiliation Laboratory of Molecular Medicine, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Yukinori Okada

Affiliations Laboratory for Statistical Analysis, Center for Genomic Medicine, Institute of Physical and Chemical Research (Center for Genomic Medicine, RIKEN), Kanagawa, Japan, Department of Allergy and Rheumatology, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Yoichiro Kamatani

Affiliation Centre d’Etude du Polymorphisme Humain, Paris, France

Michiaki Kubo

Affiliation Laboratory for Genotyping Development, Center for Genomic Medicine, RIKEN, Kanagawa, Japan

Koichi Matsuda

Affiliation Laboratory of Molecular Medicine, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Yusuke Nakamura

yusuke@ims.u-tokyo.ac.jp

Affiliation Laboratory of Molecular Medicine, Institute of Medical Science, The University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN KM YO YK MK WO. Performed the experiments: WO MK. Analyzed the data: YO YK WO. Contributed reagents/materials/analysis tools: YN KM YO MK. Wrote the paper: WO YO YN. Summarized the whole results: WO.