About the Authors

Long R. Jiao

Affiliation Hepato-Pancreato-Biliary Surgical Unit, Department of Surgery and Cancer, Imperial College, Hammersmith Hospital, London, UK

Adam E. Frampton

Affiliation Hepato-Pancreato-Biliary Surgical Unit, Department of Surgery and Cancer, Imperial College, Hammersmith Hospital, London, UK

Jimmy Jacob

Affiliation Division of Oncology, Department of Surgery and Cancer, MRC Cyclotron Building, Imperial College, Hammersmith Hospital, London, UK

Loredana Pellegrino

Affiliation Division of Oncology, Department of Surgery and Cancer, MRC Cyclotron Building, Imperial College, Hammersmith Hospital, London, UK

Jonathan Krell

Affiliation Division of Oncology, Department of Surgery and Cancer, MRC Cyclotron Building, Imperial College, Hammersmith Hospital, London, UK

Georgios Giamas

Affiliation Division of Oncology, Department of Surgery and Cancer, MRC Cyclotron Building, Imperial College, Hammersmith Hospital, London, UK

Nicole Tsim

Affiliation Hepato-Pancreato-Biliary Surgical Unit, Department of Surgery and Cancer, Imperial College, Hammersmith Hospital, London, UK

Panagiotis Vlavianos

Affiliation Department of Gastroenterology, Imperial College NHS Trust, Hammersmith Hospital, London, UK

Patrizia Cohen

Affiliation Department of Histopathology, Imperial College NHS Trust, Hammersmith Hospital, London, UK

Raida Ahmad

Affiliation Department of Histopathology, Imperial College NHS Trust, Hammersmith Hospital, London, UK

Andreas Keller

Affiliation Department of Human Genetics, Saarland University, Homburg, Saar, Germany

Nagy A. Habib

Affiliation Hepato-Pancreato-Biliary Surgical Unit, Department of Surgery and Cancer, Imperial College, Hammersmith Hospital, London, UK

Justin Stebbing

Affiliation Division of Oncology, Department of Surgery and Cancer, MRC Cyclotron Building, Imperial College, Hammersmith Hospital, London, UK

Leandro Castellano

l.castellano@imperial.ac.uk

Affiliation Division of Oncology, Department of Surgery and Cancer, MRC Cyclotron Building, Imperial College, Hammersmith Hospital, London, UK

Competing Interests

Dr. Andreas Keller was affiliated with Febit Biomed GMBH at the time of the study. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials. The other authors have declared that on competing interests exist.

Author Contributions

Conceived and designed the experiments: LRJ AF JS LC. Performed the experiments: LRJ AEF JJ LP JK LC RA PC. Analyzed the data: AEF LRJ JK JS AK LC RA PC. Contributed reagents/materials/analysis tools: LRJ NT NAH RA PC. Wrote the paper: LC AEF LRJ JS RA. Critical revision of the manuscript for important intellectual content: LRJ AEF JJ LP GG PV NAH JS LC.