About the Authors

Judith G. M. Bergboer

Affiliation Department of Dermatology, Radboud University Nijmegen Medical Centre, Nijmegen Centre for Molecular Life Sciences, Nijmegen, The Netherlands

Maša Umićević-Mirkov

Affiliation Department of Human Genetics, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Jaap Fransen

Affiliation Department of Rheumatology, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Martin den Heijer

Affiliation Department of Endocrinology and Department of Epidemiology and Biostatistics, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Barbara Franke

Affiliation Department of Human Genetics, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Piet L. C. M. van Riel

Affiliation Department of Rheumatology, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Joost Schalkwijk

Affiliation Department of Dermatology, Radboud University Nijmegen Medical Centre, Nijmegen Centre for Molecular Life Sciences, Nijmegen, The Netherlands

Marieke J. H. Coenen

m.coenen@antrg.umcn.nl

Affiliation Department of Human Genetics, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

on behalf of the Nijmegen Biomedical Study principal investigators

A list of principal investigators of the Nijmegen Biomedical Study is provided in the Acknowledgments.

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JB MU JF JS MC. Performed the experiments: JB MU. Analyzed the data: JB MU JF MC. Contributed reagents/materials/analysis tools: MH BF PR JS MC. Wrote the paper: JB MU MC JS. Revised the paper: JF MH BF PR. Final approval: JB MU JF MH BF PR JS MC.