About the Authors

Viviane Hénaux

henaux@wisc.edu

Affiliation Department of Forest and Wildlife Ecology, University of Wisconsin, Madison, Wisconsin, United States of America

Michael D. Samuel

Affiliation Wisconsin Cooperative Wildlife Research Unit, U.S. Geological Survey, Madison, Wisconsin, United States of America

Robert J. Dusek

Affiliation National Wildlife Health Center, U.S. Geological Survey, Madison, Wisconsin, United States of America

Joseph P. Fleskes

Affiliation Western Ecological Research Center, U.S. Geological Survey, Dixon, California, United States of America

Hon S. Ip

Affiliation National Wildlife Health Center, U.S. Geological Survey, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VH MDS RJD JPF HSI. Performed the experiments: VH MDS RJD JPF HSI. Analyzed the data: VH MDS. Contributed reagents/materials/analysis tools: VH MDS RJD JPF HSI. Wrote the paper: VH MDS RJD JPF HSI.