About the Authors

Sarah N. Bevins

bevins@rams.colostate.edu

Affiliation USDA National Wildlife Disease Program, Fort Collins, Colorado, United States of America

Scott Carver

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Erin E. Boydston

Affiliation U.S. Geological Survey, Western Ecological Research Center, Thousand Oaks, California, United States of America

Lisa M. Lyren

Affiliation U.S. Geological Survey, Western Ecological Research Center, Thousand Oaks, California, United States of America

Mat Alldredge

Affiliation Colorado Parks and Wildlife, Fort Collins, Colorado, United States of America

Kenneth A. Logan

Affiliation Colorado Parks and Wildlife, Montrose, Colorado, United States of America

Seth P. D. Riley

Affiliation National Park Service, Thousand Oaks, California, United States of America

Robert N. Fisher

Affiliation U.S. Geological Survey, Western Ecological Research Center, San Diego, California, United States of America

T. Winston Vickers

Affiliation Wildlife Health Center, University of California Davis, Davis, California, United States of America

Walter Boyce

Affiliation Wildlife Health Center, University of California Davis, Davis, California, United States of America

Mo Salman

Affiliation Department of Clinical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Michael R. Lappin

Affiliation Department of Clinical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Kevin R. Crooks

Affiliation Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America

Sue VandeWoude

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ML KC SV. Performed the experiments: SB. Analyzed the data: SB. Contributed reagents/materials/analysis tools: EB LL MA KL SR RF TV WB MS ML. Wrote the paper: SB SC MS KC SV.