About the Authors
- Raymond Alfred
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* E-mail: raymond_alfred@yahoo.com
Affiliations Borneo Species Programme, World Wildlife Fund-Malaysia, Kota Kinabalu, Sabah, Malaysia, Borneo Conservation Trust, Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia, Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Sabah, Malaysia
- Abd Hamid Ahmad
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Affiliation Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Sabah, Malaysia
- Junaidi Payne
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Affiliation Borneo Rhino Alliance (BORA), Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu, Sabah, Malaysia
- Christy Williams
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Affiliation World Wildlife Fund Asian Rhinoceros and Elephant Action Strategy, World Wildlife Fund-International, Gland, Switzerland
- Laurentius Nayan Ambu
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Affiliation Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia
- Phua Mui How
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Affiliation School of International Forestry, Universiti Malaysia Sabah, Kota Kinabalu, Sabah, Malaysia
- Benoit Goossens
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Affiliations Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia, Danau Girang Field Centre, Sabah Wildlife Department, Kota Kinabalu, Sabah, Malaysia, Organisms and Environment Division, School of Biosciences, Cardiff University, Cardiff, United ingdom
Competing Interests
The authors have declared that no competing interests exist.
Author Contributions
Conceived and designed the experiments: RA AHA JP CW LN PMH BG. Performed the experiments: RA. Analyzed the data: RA JP CW BG. Wrote the paper: RA BG.