About the Authors

Natacha Scarafone

Affiliation Laboratory of Enzymology and Protein Folding, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

Coralie Pain

Affiliation Laboratory of Enzymology and Protein Folding, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

Anthony Fratamico

Current address: Laboratory of Plant Biochemistry and Photobiology, University of Liège, Liège, Belgium

Affiliation Laboratory of Enzymology and Protein Folding, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

Gilles Gaspard

Current address: ProGenosis S.A., Sart-Tilman, Liège, Belgium

Affiliation Biological Macromolecules, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

Nursel Yilmaz

Current address: ProGenosis S.A., Sart-Tilman, Liège, Belgium

Affiliation Biological Macromolecules, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

Patrice Filée

Current address: ProGenosis S.A., Sart-Tilman, Liège, Belgium

Affiliation Biological Macromolecules, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

Moreno Galleni

Affiliation Biological Macromolecules, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

André Matagne

Affiliation Laboratory of Enzymology and Protein Folding, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

Mireille Dumoulin

mdumoulin@ulg.ac.be

Affiliation Laboratory of Enzymology and Protein Folding, Centre for Protein Engineering, Institute of Chemistry, University of Liège, Liège, Belgium

Competing Interests

The authors have read the journal's policy and have the following conflicts: Current address of GG, NY and PF: ProGenosis S.A., Boulevard du Rectorat, 27b, Sart-Tilman, 4000 Liège, Belgium. These three persons are employed by the company. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NS AM MD. Performed the experiments: NS CP AF. Analyzed the data: NS CP AM MD. Contributed reagents/materials/analysis tools: NY GG PF MG. Wrote the paper: NS AM MD.