About the Authors

Israt Jahan

Affiliation University of Iowa, Department of Biology, Iowa City, Iowa, United States of America

Ning Pan

Affiliation University of Iowa, Department of Biology, Iowa City, Iowa, United States of America

Jennifer Kersigo

Affiliation University of Iowa, Department of Biology, Iowa City, Iowa, United States of America

Lilian E. Calisto

Affiliation Creighton University, Department of Biomedical Sciences, Omaha, Nebraska, United States of America

Ken A. Morris

Affiliation Creighton University, Department of Biomedical Sciences, Omaha, Nebraska, United States of America

Benjamin Kopecky

Affiliation University of Iowa, Department of Biology, Iowa City, Iowa, United States of America

Jeremy S. Duncan

Affiliation University of Iowa, Department of Biology, Iowa City, Iowa, United States of America

Kirk W. Beisel

Contributed equally to this work with: Kirk W. Beisel, Bernd Fritzsch

Affiliation Creighton University, Department of Biomedical Sciences, Omaha, Nebraska, United States of America

Bernd Fritzsch

Contributed equally to this work with: Kirk W. Beisel, Bernd Fritzsch

bernd-fritzsch@uiowa.edu

Affiliation University of Iowa, Department of Biology, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IJ KWB BF. Performed the experiments: IJ NP JK LEC KAM BK JSD BF. Analyzed the data: IJ BF. Contributed reagents/materials/analysis tools: IJ NP JK. Wrote the paper: IJ KWB BF.