About the Authors

Markus Horning

markus.horning@oregonstate.edu

Affiliations Department of Fisheries and Wildlife, Marine Mammal Institute, Oregon State University, Newport, Oregon, United States of America, Coastal Oregon Marine Experiment Station, Newport, Oregon, United States of America

Jo-Ann E. Mellish

Affiliations Alaska SeaLife Center, Seward, Alaska, United States of America, School of Fisheries and Ocean Sciences, University of Alaska Fairbanks, Alaska, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MH. Performed the experiments: MH JEM. Analyzed the data: MH. Contributed reagents/materials/analysis tools: MH JEM. Wrote the paper: MH. Conducted field work: JEM MH Obtained research permits and supervised husbandry: JEM Conducted physiological control studies: JEM Revised and approved final version of paper: MH JEM.