About the Authors

Byoungkoo Lee

Affiliation Department of Biological Sciences and Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Philip R. LeDuc

Affiliation Departments of Mechanical and Biomedical Engineering, and Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Russell Schwartz

russells@andrew.cmu.edu

Affiliation Department of Biological Sciences and Lane Center for Computational Biology, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BL PRL RS. Performed the experiments: BL. Analyzed the data: BL PRL RS. Contributed reagents/materials/analysis tools: BL. Wrote the paper: BL PRL RS. Designed and implemented simulation software: BL.