About the Authors

Ian Holmes

Affiliation Department of Bioengineering, University of California, Berkeley, California, United States of America

Keith Harris

Affiliation School of Engineering, University of Glasgow, Glasgow, United Kingdom

Christopher Quince

christopher.quince@glasgow.ac.uk

Affiliation School of Engineering, University of Glasgow, Glasgow, United Kingdom

Competing Interests

KH is directly funded through a Unilever research grant to develop bioinformatics tools. All tools developed under this grant are being released open source. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: IH KH CQ. Performed the experiments: IH KH CQ. Analyzed the data: IH KH CQ. Contributed reagents/materials/analysis tools: IH KH CQ. Wrote the paper: IH KH CQ.