About the Authors

Joshua T. Selsby

Affiliations Department of Physiology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Animal Science, College of Agriculture and Life Sciences, Iowa State University, Ames, Iowa, United States of America

Kevin J. Morine

Affiliation Department of Physiology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Klara Pendrak

Affiliation Department of Physiology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Elisabeth R. Barton

Affiliation Department of Anatomy and Cell Biology, School of Dental Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

H. Lee Sweeney

lsweeney@mail.med.upenn.edu

Affiliation Department of Physiology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JTS ERB HLS. Performed the experiments: JTS KJM KP. Analyzed the data: JTS KJM ERB KP HLS. Contributed reagents/materials/analysis tools: ERB HLS. Wrote the paper: JTS HLS. Critical evaluation of manuscript: KJM ERB KP.