About the Authors

Valentin Fischer

v.fischer@ulg.ac.be

Affiliations Geology department, University of Liège, Liège, Belgium, Paleontology Department, Royal Belgian Institute of Natural Sciences, Brussels, Belgium

Michael W. Maisch

Current address: Tübingen, Germany

Darren Naish

Affiliations School of Earth and Environmental Sciences, University of Portsmouth, Portsmouth, United Kingdom, School of Ocean and Earth Sciences, University of Southampton, Southampton, United Kingdom

Ralf Kosma

Affiliation Staatliches Naturhistorisches Museum, Braunschweig, Germany

Jeff Liston

Current address: National Museums Scotland, Edinburgh, Scotland

Affiliation Division of Environmental and Evolutionary Biology, School of Life Sciences, College of Medical Veterinary and Life Sciences, University of Glasgow, Glasgow, Scotland

Ulrich Joger

Affiliation Staatliches Naturhistorisches Museum, Braunschweig, Germany

Fritz J. Krüger

Affiliation Staatliches Naturhistorisches Museum, Braunschweig, Germany

Judith Pardo Pérez

Affiliation Institut für Geowissenschaften, Ruprecht-Karls-Universität Heidelberg, Heidelberg, Germany

Jessica Tainsh

Affiliation Division of Environmental and Evolutionary Biology, School of Life Sciences, College of Medical Veterinary and Life Sciences, University of Glasgow, Glasgow, Scotland

Robert M. Appleby

† Deceased.

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VF. Performed the experiments: VF. Analyzed the data: VF. Contributed reagents/materials/analysis tools: UJ RK FJK. Wrote the paper: VF DN JL RK. Described the Cremlingen specimen: VF MWM JPP. Described the Speeton specimen: VF RMA JT. Described the CAMSM and BMNH specimens: VF.