About the Authors

David W. Johnston

david.johnston@duke.edu

Affiliation Duke University Marine Laboratory, Division of Marine Science and Conservation, Nicholas School of the Environment, Duke University, Beaufort, North Carolina, United States of America

Matthew T. Bowers

Affiliation Duke University Marine Laboratory, Division of Marine Science and Conservation, Nicholas School of the Environment, Duke University, Beaufort, North Carolina, United States of America

Ari S. Friedlaender

Affiliation Duke University Marine Laboratory, Division of Marine Science and Conservation, Nicholas School of the Environment, Duke University, Beaufort, North Carolina, United States of America

David M. Lavigne

Affiliation International Fund for Animal Welfare, Guelph, Ontario, Canada

Competing Interests

Author DML is the Scientific Advisor for the International Fund for Animal Welfare, the organization which helped fund this research. This does not alter the authors′ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: DWJ ASF DML. Performed the experiments: DWJ ASF MTB. Analyzed the data: DWJ MTB. Wrote the paper: DWJ MTB ASF DML.