About the Authors

Sadie J. Ryan

sjryan@esf.edu

Affiliation Department of Environmental and Forest Biology, SUNY College of Environmental Science and Forestry, Syracuse, New York, United States of America

Peter D. Walsh

Affiliation Department of Archaeology and Anthropology, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: SJR PDW. Contributed reagents/materials/analysis tools: SJR PDW. Wrote the paper: SJR PDW.