About the Authors

Philip Riordan

philip.riordan@zoo.ox.ac.uk

Affiliation Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Oxford, United Kingdom

Richard John Delahay

Affiliation The Food and Environment Research Agency, York, United Kingdom

Chris Cheeseman

Affiliation The Food and Environment Research Agency, York, United Kingdom

Paul James Johnson

Affiliation Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Oxford, United Kingdom

David Whyte Macdonald

Affiliation Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PR RJD CC DWM. Performed the experiments: PR RJD. Analyzed the data: PR PJ. Contributed reagents/materials/analysis tools: RJD CC. Wrote the paper: PR RJD CC PJ DWM.