About the Authors

Mark A. Little

Contributed equally to this work with: Mark A. Little, Bahjat Al-Ani, Shuyu Ren

jeremysd@u.washington.edu (JSD) ; mark.little@ucl.ac.uk (ML)

Affiliation Centre for Nephrology, Royal Free Hospital, University College London, London, United Kingdom

Bahjat Al-Ani

Contributed equally to this work with: Mark A. Little, Bahjat Al-Ani, Shuyu Ren

Current address: Department of Physiology, College of Medicine, King Khalid University, Abha, Saudi Arabia

Affiliation Renal Institute of Birmingham, School of Infection, Immunology and Inflammation, University of Birmingham, Birmingham, United Kingdom

Shuyu Ren

Contributed equally to this work with: Mark A. Little, Bahjat Al-Ani, Shuyu Ren

Affiliation Division of Nephrology, Department of Medicine, Center for Lung Biology and Institute for Stem Cell and Regenerative Medicine, University of Washington, Seattle, Washington, United States of America

Hamad Al-Nuaimi

Affiliation Renal Institute of Birmingham, School of Infection, Immunology and Inflammation, University of Birmingham, Birmingham, United Kingdom

Maurilo Leite Jr

Affiliations Division of Nephrology, Department of Medicine, Center for Lung Biology and Institute for Stem Cell and Regenerative Medicine, University of Washington, Seattle, Washington, United States of America, Division of Nephrology, Federal University of Rio de Janeiro, Rio de Janeiro, Brazil

Charles E. Alpers

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

Caroline O. Savage

Affiliations Renal Institute of Birmingham, School of Infection, Immunology and Inflammation, University of Birmingham, Birmingham, United Kingdom, GlaxoSmithKline UK Ltd, Uxbridge, United Kingdom

Jeremy S. Duffield

jeremysd@u.washington.edu (JSD) ; mark.little@ucl.ac.uk (ML)

Affiliation Division of Nephrology, Department of Medicine, Center for Lung Biology and Institute for Stem Cell and Regenerative Medicine, University of Washington, Seattle, Washington, United States of America

Competing Interests

C.O.S. is employed by GSK. This study was partly funded by a GRIP award from Genzyme. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MAL COS JSD. Performed the experiments: BA-A SR HA-N ML MAL JSD. Analyzed the data: MAL COS CEA JSD. Wrote the manuscript: MAL COS JSD.