About the Authors
- Takeshi Sato
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Contributed equally to this work with: Takeshi Sato, Yoshimi Aiyama, Mayuko Ishii-Inagaki
Affiliations Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan, Drug Safety Research Labs, Astellas Pharma Inc., Osaka, Japan
- Yoshimi Aiyama
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Contributed equally to this work with: Takeshi Sato, Yoshimi Aiyama, Mayuko Ishii-Inagaki
Affiliation Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan
- Mayuko Ishii-Inagaki
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Contributed equally to this work with: Takeshi Sato, Yoshimi Aiyama, Mayuko Ishii-Inagaki
Affiliation Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan
- Kenshiro Hara
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Affiliation Division of Germ Cell Biology, National Institute for Basic Biology and Department of Basic Biology, School of Life Science, Graduate University for Advanced Studies (SOKENDAI), Okazaki, Japan
- Naoki Tsunekawa
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Affiliation Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan
- Kyoko Harikae
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Affiliation Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan
- Mami Uemura-Kamata
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Affiliations Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan, Center for Experimental Animal, Tokyo Medical and Dental University, Tokyo, Japan
- Mai Shinomura
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Affiliation Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan
- Xiao Bo Zhu
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Affiliation Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan
- Seishi Maeda
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Affiliation Division of Cell Biology, Department of Anatomy, Hyogo College of Medicine, Nishinomiya, Japan
- Sachi Kuwahara-Otani
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Affiliation Division of Cell Biology, Department of Anatomy, Hyogo College of Medicine, Nishinomiya, Japan
- Akihiko Kudo
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Affiliation Department of Anatomy, Kyorin University School of Medicine, Mitaka, Japan
- Hayato Kawakami
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Affiliation Department of Anatomy, Kyorin University School of Medicine, Mitaka, Japan
- Masami Kanai-Azuma
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Affiliation Center for Experimental Animal, Tokyo Medical and Dental University, Tokyo, Japan
- Michio Fujiwara
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Affiliation Drug Safety Research Labs, Astellas Pharma Inc., Osaka, Japan
- Yoichi Miyamae
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Affiliation Drug Safety Research Labs, Astellas Pharma Inc., Osaka, Japan
- Shosei Yoshida
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Affiliation Division of Germ Cell Biology, National Institute for Basic Biology and Department of Basic Biology, School of Life Science, Graduate University for Advanced Studies (SOKENDAI), Okazaki, Japan
- Makoto Seki
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Affiliation Division of Cell Biology, Department of Anatomy, Hyogo College of Medicine, Nishinomiya, Japan
- Masamichi Kurohmaru
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Affiliation Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan
- Yoshiakira Kanai
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* E-mail: aykanai@mail.ecc.u-tokyo.ac.jp
Affiliation Department of Veterinary Anatomy, The University of Tokyo, Tokyo, Japan
Competing Interests
TS, MF, and YM are employed by Astellas Pharma Inc. However, this does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.
Author Contributions
Conceived and designed the experiments: TS YA MI-I YK. Performed the experiments: TS YA MI-I K. Hara NT K. Harikae MU-K M. Shinomura XBZ SM SK-O AK H. Kawakami MK-A YM. Analyzed the data: TS YA MI-I K. Hara NT K. Harikae MU-K M. Shinomura XBZ M. Seki SM SK-O AK H. Kawakami MK-A YM. Contributed reagents/materials/analysis tools: K. Hara SY. Wrote the paper: SY MK YK. Permitted and indirectly supported Dr. Sato's (TS) work: MF YM.