About the Authors

Heleen C. Druce

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, Westville Campus, University of KwaZulu-Natal, Durban, South Africa

Robin L. Mackey

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, Westville Campus, University of KwaZulu-Natal, Durban, South Africa

Rob Slotow

slotow@ukzn.ac.za

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, Westville Campus, University of KwaZulu-Natal, Durban, South Africa

Competing Interests

The authors have read the journal's policy and have the following conflicts: Distell (PTY) Ltd made a donation to UKZN for research on elephant conservation. This is a social responsibility donation, and is not their core business. There are no restrictions on publication from this work. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HCD RLM RS. Performed the experiments: HCD. Analyzed the data: HCD RLM RS. Contributed reagents/materials/analysis tools: RLM RS. Wrote the paper: HCD. Contributed editorial improvements to paper: RLM RS.