About the Authors

Emma R. Travis

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

William H. Gaze

Contributed equally to this work with: William H. Gaze, Alessandra Pontiroli

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

Alessandra Pontiroli

Contributed equally to this work with: William H. Gaze, Alessandra Pontiroli

A.Pontiroli@warwick.ac.uk

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

Francis P. Sweeney

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

David Porter

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

Sam Mason

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

Matthew J. C. Keeling

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

Rebecca M. Jones

Affiliation Animal Health and Veterinary Laboratories Agency, Weybridge, United Kingdom

Jason Sawyer

Affiliation Animal Health and Veterinary Laboratories Agency, Weybridge, United Kingdom

Alicia Aranaz

Affiliation School of Veterinary Sciences. Centro de Vigilancia Sanitaria Veterinaria (VISAVET) and Animal Health Department. Universidad Complutense de Madrid, Madrid, Spain

Elena Castellanos Rizaldos

Affiliation School of Veterinary Sciences. Centro de Vigilancia Sanitaria Veterinaria (VISAVET) and Animal Health Department. Universidad Complutense de Madrid, Madrid, Spain

Jennifer Cork

Affiliation Animal Health and Veterinary Laboratories Agency, Weybridge, United Kingdom

Richard J. Delahay

Affiliation Food and Environment Research Agency, Sand Hutton, York, United Kingdom

Gavin J. Wilson

Affiliation Food and Environment Research Agency, Sand Hutton, York, United Kingdom

R. Glyn Hewinson

These authors also contributed equally to this work.

Affiliation Animal Health and Veterinary Laboratories Agency, Weybridge, United Kingdom

Orin Courtenay

These authors also contributed equally to this work.

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

Elizabeth M. H. Wellington

These authors also contributed equally to this work.

Affiliation University of Warwick, School of Life Sciences, Coventry, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ERT WG AP OC EMW RGH. Performed the experiments: FPS ECR JC DP. Analyzed the data: AP SM ERT. Contributed reagents/materials/analysis tools: AP FPS DP SM MJCK RJD GJW RMJ JS AA. Wrote the paper: ERT AP.