About the Authors

Giada Cordoni

Affiliations Natural History Museum, University of Pisa, Pisa, Italy, Vademecos, Biology Professional Association, Viareggio, Italy

Elisabetta Palagi

betta.palagi@museo.unipi.it

Affiliations Natural History Museum, University of Pisa, Pisa, Italy, Unit of Cognitive Primatology and Primate Center, Institute of Cognitive Sciences and Technologies, Consiglio Nazionale delle Ricerche (CNR), Roma, Italy

Competing Interests

ZooParc de Beauval and Dierenpark Amersfoort were funders of this study. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GC EP. Performed the experiments: GC EP. Analyzed the data: GC EP. Contributed reagents/materials/analysis tools: GC EP. Wrote the paper: GC EP.