About the Authors

Wai-Hong Wu

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Elizabeth Gersch

Affiliation Department of Molecular, Cellular and Developmental Biology, University of Colorado, Boulder, Colorado, United States of America

Kihyuck Kwak

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Subhashini Jagu

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Balasubramanyam Karanam

Affiliation Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America

Warner K. Huh

Affiliation Department of Gynecologic Oncology, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Robert L. Garcea

Affiliation Department of Molecular, Cellular and Developmental Biology, University of Colorado, Boulder, Colorado, United States of America

Richard B. S. Roden

roden@jhmi.edu

Affiliations Department of Pathology, Johns Hopkins University, Baltimore, Maryland, United States of America, Department of Oncology, Johns Hopkins University, Baltimore, Maryland, United States of America, Department of Gynecology and Obstetrics, Johns Hopkins University, Baltimore, Maryland, United States of America

Competing Interests

SJ and RBSR are co-inventors on L2 patents licensed to Shantha Biotechnics Ltd., GlaxoSmithKline, PaxVax, Inc. and Acambis, Inc. The terms of these arrangements are managed by Johns Hopkins University in accordance with its conflict of interest policies. RLG has intellectual property concerning capsomer vaccines, managed by the University of Colorado Technology Transfer Office. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RBR RLG WKH. Performed the experiments: W-HW EG KK SJ BK. Analyzed the data: W-HW EG KK SJ BK RBR RLG WKH. Contributed reagents/materials/analysis tools: KK SJ BK. Wrote the paper: RBR RLG WKH.