About the Authors

Indrani Mukhopadhya

Contributed equally to this work with: Indrani Mukhopadhya, Richard Hansen

Affiliation Gastrointestinal Research Group, Division of Applied Medicine, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

Richard Hansen

Contributed equally to this work with: Indrani Mukhopadhya, Richard Hansen

Affiliations Gastrointestinal Research Group, Division of Applied Medicine, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom, Child Health, University of Aberdeen, Royal Aberdeen Children's Hospital, Foresterhill, Aberdeen, United Kingdom

Charlotte E. Nicholl

Affiliation Gastrointestinal Research Group, Division of Applied Medicine, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

Yazeid A. Alhaidan

Affiliation Gastrointestinal Research Group, Division of Applied Medicine, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

John M. Thomson

Affiliation Gastrointestinal Research Group, Division of Applied Medicine, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

Susan H. Berry

Affiliation Gastrointestinal Research Group, Division of Applied Medicine, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

Craig Pattinson

Affiliation Aberdeen Proteomics Group, Institute of Medical Sciences, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

David A. Stead

Affiliation Aberdeen Proteomics Group, Institute of Medical Sciences, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

Richard K. Russell

Affiliation Department of Paediatric Gastroenterology, Royal Hospital for Sick Children, Glasgow, United Kingdom

Emad M. El-Omar

Affiliation Gastrointestinal Research Group, Division of Applied Medicine, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

Georgina L. Hold

g.l.hold@abdn.ac.uk.

Affiliation Gastrointestinal Research Group, Division of Applied Medicine, University of Aberdeen, Foresterhill, Aberdeen, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IM RH EME-O GLH. Performed the experiments: IM RH CEN YAA SHB CP DAS. Analyzed the data: IM RH JMT RKR GLH. Contributed reagents/materials/analysis tools: JMT RKR EME-O. Wrote the paper: IM RH GLH.