About the Authors

Shinya Sato

s.sato@rbge.ac.uk

Affiliations Royal Botanic Garden Edinburgh, Edinburgh, United Kingdom, School of Biological Sciences, Institute of Molecular Plant Sciences, University of Edinburgh, Edinburgh, United Kingdom

Gordon Beakes

Affiliation School of Biology, Newcastle University, Newcastle upon Tyne, United Kingdom

Masahiko Idei

Affiliation Department of Biology, Bunkyo University, Koshigaya, Saitama, Japan

Tamotsu Nagumo

Affiliation Department of Biology, The Nippon Dental University, Chiyoda-ku, Tokyo, Japan

David G. Mann

Affiliation Royal Botanic Garden Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS DGM. Performed the experiments: SS. Analyzed the data: SS GB MI TN DGM. Contributed reagents/materials/analysis tools: SS GB MI TN DGM. Wrote the paper: SS DGM.