About the Authors

James F. Graham

Affiliation The Roslin Institute and R(D)SVS, Neuropathogenesis Division, University of Edinburgh, Easter Bush, Roslin, Edinburgh, Midlothian, United Kingdom

Dominic Kurian

Affiliation Institute for Animal Health, Compton, Newbury, Berkshire, United Kingdom

Sonya Agarwal

Affiliation The Roslin Institute and R(D)SVS, Neuropathogenesis Division, University of Edinburgh, Easter Bush, Roslin, Edinburgh, Midlothian, United Kingdom

Lorna Toovey

Affiliation The Roslin Institute and R(D)SVS, Neuropathogenesis Division, University of Edinburgh, Easter Bush, Roslin, Edinburgh, Midlothian, United Kingdom

Lawrence Hunt

Affiliation Institute for Animal Health, Compton, Newbury, Berkshire, United Kingdom

Louise Kirby

Affiliation The Roslin Institute and R(D)SVS, Neuropathogenesis Division, University of Edinburgh, Easter Bush, Roslin, Edinburgh, Midlothian, United Kingdom

Teresa J. T. Pinheiro

Affiliation School of Biological Sciences, University of Warwick, Coventry, United Kingdom

Steven J. Banner

Affiliation The Roslin Institute and R(D)SVS, Neuropathogenesis Division, University of Edinburgh, Easter Bush, Roslin, Edinburgh, Midlothian, United Kingdom

Andrew C. Gill

andrew.gill@roslin.ed.ac.uk

Affiliation The Roslin Institute and R(D)SVS, Neuropathogenesis Division, University of Edinburgh, Easter Bush, Roslin, Edinburgh, Midlothian, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LH LK TJTP SJB ACG. Performed the experiments: JFG DK SA LT. Analyzed the data: JFG DK ACG. Wrote the paper: JFG ACG.