About the Authors

Richard Francis

Affiliation Department of Developmental Biology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Xin Xu

Affiliation Genetics and Development Biology Center, National Heart Lung and Blood Institute, Bethesda, Maryland, United States of America

Hyunsoo Park

Affiliation Genetics and Development Biology Center, National Heart Lung and Blood Institute, Bethesda, Maryland, United States of America

Chin-Jen Wei

Affiliation Genetics and Development Biology Center, National Heart Lung and Blood Institute, Bethesda, Maryland, United States of America

Stephen Chang

Affiliation Genetics and Development Biology Center, National Heart Lung and Blood Institute, Bethesda, Maryland, United States of America

Bishwanath Chatterjee

Affiliation Genetics and Development Biology Center, National Heart Lung and Blood Institute, Bethesda, Maryland, United States of America

Cecilia Lo

cel36@pitt.edu

Affiliation Department of Developmental Biology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RF XX CJW BC CL. Performed the experiments: RF XX HP CJW SC BC. Analyzed the data: RF XX HP CL. Contributed reagents/materials/analysis tools: BC CL. Wrote the paper: RF CL.